B.Sc. Microbiology, Internat. Agricultural College Larenstein, Wageningen, NL., 1992.
M.Sc. Cellular Biology and Microbial Ecology, University of Wageningen, NL., 1996.
Ph.D. Molecular Microbial Ecology, Institute for the Chemistry and Biology of the Marine Environment (ICBM), University of Oldenburg, Germany, 2000
Survival of nucleic acids in Quaternary lacustrine and marine sediments
Developing new molecular tools for paleogenetics
Reconstruction of the ancient lacustrine- and marine microbial diversity, and hence, reconstruction of climate-induced changes of paleoenvironments based on the combined stratigraphic analysis of geolipid biomarkers and preserved fossil ribosomal RNA encoding genes
Reconstruction of ancient and contemporary metabolic processes in lacustrine and marine environments as well as the prokaryotes involved in these processes based on the analysis of modern and preserved functional genes and gene products
Microbial diversity and processes in stratified environments