B.A. Colgate University, 1982, Geology
M.S. University of California, Davis, 1984, Geology
Ph.D. Scripps Institution of Oceanography, 1990, Oceanography
PostDoc, Wadsworth Center, NY State Department of Health, 1990-1994, Cell Biology
Fulbright Scholar, University of Oslo, Department of Geology, 1992, Foraminiferal ecology
Benthic foraminiferal ecology and paleoecology; Biogeochemistry of redox boundary sediments in both modern and ancient environments; Microaerophilic and anaerobic protists from Oxygen Minimum Zones and methane and cold seeps; Protistan-prokaryote symbioses; Culturing of deep-sea benthic foraminifera to ground truth paleoceanographic proxies; Microbial communities of laminated sediments; Sub-millimeter life positions of microorganisms in sediments; Application of various microscopic techniques (TEM, SEM, LSCM) to marine microbial ecology.