Emeritus Research Scholar
Office Phone: +1 508 289 2751
Fax: +1 508 457 2134
WHOI Mailing Address:
Woods Hole Oceanographic Institution
266 Woods Hole Rd.
Woods Hole, MA 02543-1050
B.S. Michigan State University, 1971, Zoology
Ph.D. Michigan State University, 1974, Zoology
Mathematical population and community ecology; matrix population models for organisms with complex life cycles; life history theory; recruitment; nonlinear dynamics and chaos; population-level ecotoxicology; conservation biology; marine mammal and seabird populations.